1Department of Anatomy, Cell Biology & Physiology, Indiana University School of Medicine, Indianapolis, Indiana, USA
2Department of Orthopaedic Surgery, School of Medicine, UC Davis Health, Sacramento, California, USA
3Richard L. Roudebush Veterans Affairs Medical Center, Indianapolis, Indiana, USA
4Department of Biomedical Engineering, Indiana University–Purdue University at Indianapolis, Indianapolis, Indiana, USA
5Indiana Center for Musculoskeletal Health, Indianapolis, Indiana, USA